Jeremy N. Morris, a British epidemiologist whose comparison of heart-attack rates among double-decker bus drivers and conductors in London in the late 1940s and early '50s laid the scientific groundwork for the modern aerobics movement, died Oct. 28 in Hampstead, London. He was 99 ½.

"He always insisted on adding the ½," said his daughter, Julie Zalewska.

The cause of death, she said, was pneumonia and kidney failure.

It had long been surmised that exercise and a healthy heart were correlated.

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